Bioluminescence is the production and emission of light by a living organism. It is a form of chemiluminescence. Bioluminescence occurs widely in marine vertebrates and invertebrates, as well as in some fungi, microorganisms including some bioluminescent bacteria, and terrestrial arthropods such as fireflies. In some animals, the light is bacteriogenic, produced by symbiotic bacteria such as those from the genus Vibrio; in others, it is autogenic, produced by the animals themselves.
In a general sense, the principal chemical reaction in bioluminescence involves a light-emitting molecule and an enzyme, generally called luciferin and luciferase, respectively. Because these are generic names, luciferins and luciferases are often distinguished by the species or group, e.g. firefly luciferin. In all characterized cases, the enzyme catalyzes the oxidation of the luciferin.
In some species, the luciferase requires other cofactors, such as calcium or magnesium ions, and sometimes also the energy-carrying molecule adenosine triphosphate (ATP). In evolution, luciferins vary little: one in particular, coelenterazine, is found in 11 different animal phyla, though in some of these, the animals obtain it through their diet. Conversely, luciferases vary widely between different species, which is evidence that bioluminescence has arisen over 40 times in evolutionary history.
Both Aristotle and Pliny the Elder mentioned that damp wood sometimes gives off a glow. Many centuries later Robert Boyle showed that oxygen was involved in the process, in both wood and glowworms. It was not until the late nineteenth century that bioluminescence was properly investigated. The phenomenon is widely distributed among animal groups, especially in marine environments. On land it occurs in fungi, bacteria and some groups of invertebrates, including insects.
The uses of bioluminescence by animals include counterillumination camouflage, mimicry of other animals, for example to lure prey, and signaling to other individuals of the same species, such as to attract mates. In the laboratory, luciferase-based systems are used in genetic engineering and biomedical research. Researchers are also investigating the possibility of using bioluminescent systems for street and decorative lighting, and a bioluminescent plant has been created.
San Diego (/ˌsæn diˈeɪɡoʊ/ SAN dee-AY-goh, Spanish: [san ˈdjeɣo]; Spanish for ‘Saint Didacus’) is a city in the U.S. state of California on the coast of the Pacific Ocean and immediately adjacent to the Mexican border. With a 2020 population of 1,386,932, San Diego is the eighth most populous city in the United States and second most populous in California (after Los Angeles). The city is the county seat of San Diego County, the fifth most populous county in the United States, with 3,338,330 estimated residents as of 2019. The city is known for its mild year-round climate, natural deep-water harbor, extensive beaches and parks, long association with the United States Navy and Marine Corps, and recent emergence as a healthcare and biotechnology development center.
Historically home to the Kumeyaay people, San Diego is frequently referred to as the “Birthplace of California”, as it was the first site visited and settled by Europeans on what is now the West Coast of the United States. Upon landing in San Diego Bay in 1542, Juan Rodríguez Cabrillo claimed the area for Spain, forming the basis for the settlement of Alta California 200 years later. The Presidio and Mission San Diego de Alcalá, founded in 1769, formed the first European settlement in what is now California. In 1821, San Diego became part of the newly declared Mexican Empire, which reformed as the First Mexican Republic two years later. California became part of the United States in 1848 following the Mexican–American War and was admitted to the union as a state in 1850.
San Diego’s main economic engines are military and defense-related activities, tourism, international trade, research, and manufacturing. The city is the economic center of the San Diego–Tijuana conurbation, the second most populous transborder metropolitan area in the western hemisphere (after Detroit–Windsor), home to an estimated 4,922,723 people as of 2012. The primary border crossing between San Diego and Tijuana, the San Ysidro Port of Entry, is the busiest international land border crossing in the world outside of Asia (fourth-busiest overall). The city’s primary airport, San Diego International Airport, is the busiest single-runway airport in the world.[a]
Bioluminescence, San Diego museum-quality poster made on thick and durable matte paper. Add a wonderful accent to your room and office with these posters that are sure to brighten any environment.
• Paper thickness: 0.26 mm (10.3 mil)
• Paper weight: 189 g/m² (5.57 oz/y²)
• Opacity: 94%
• ISO brightness: 104%
• Giclée printing quality
• 21 × 30 cm posters are size A4
• Blank product sourced from Japan